The horse has 64 chromosomes (32 pairs), and the donkey has 62 (31 pairs). When they are mated, the mule receives 63 chromosomes (32 from the horse and 31 from the donkey). During meiosis, the chromosomes are not in pairs, and, as their number is halved, the sex cells usually end up without a complete set. This unbalanced condition does not allow the sex cells to function, resulting in sterility.
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This work is supported in part by New Technologies for Agriculture Extension grant no. 2020-41595-30123 from the USDA National Institute of Food and Agriculture. Any opinions, findings, conclusions, or recommendations expressed in this publication are those of the author(s) and do not necessarily reflect the view of the U.S. Department of Agriculture.
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